maybe a little late to the party here but i've always said "zoh-oh-zan-THELL-ay"

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As others have said, "zoo-" is two syllables, "zo-o-", both of which are originally omicron. "X" here is Greek chi, which at some point in classical Latin, by or about its transition into old Italian, begins to take up a softer "hard s" sound, found in "z" today. However, Greeks said chi as "hard k"

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Also the German pronunciation pronounces the X.

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There was a Ted talk I saw once with a marine biologist discussing the symbiotic algae and pronounced the "X", so she said "ZOOCKS-AN-Thele. And this was a marine biologist. So in the immortal words of Patrick Stewart "...eventually nobody gives a sh*t"

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The prefix "zoo-" is usually pronounced like "zoa" or "zo-o" (two separate vowel sounds next two each other) — for example, "zoospore" is pronounced "ZOH-uh-spore" — but I'm not sure about "zooxanthellae" specifically.

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